While still in the pushup position, make a fist with each hand one at a time, . This will strengthen not only your knuckles, but your entire hand.
Perform pushups, as usual, but put your weight on the first two knuckles of each hand. allowing your hands to strengthen and harden according to Wolff's law.
Make a Fist. fist stretch exercise. Hand and finger exercises can help strengthen your hands and fingers, increase your range of motion, and give you pain relief.
When lifting heavy weights, the grip is often the first thing to go. Workouts that strengthen the forearm flexors will help enhance overall grip.
Other exercises strengthen muscles around a joint to generate more power or to build greater endurance. These Make a straight fist; return to a straight hand.